Methods and tools to characterise soils for plant available water capacity (North Star)

Take home messages

  • Information regarding the plant available water (PAW) at a point in time, particularly at planting, can be useful in a range of crop management decisions. Estimating PAW, whether through use of a soil water monitoring device or a push probe, requires knowledge of the plant available water capacity (PAWC) and/or the Crop Lower Limit (CLL).
  • A wide variety of soils in the northern region have been characterised for PAWC and the characterisations are publicly available in the APSoil database, which can be viewed in Google Earth and in the ‘SoilMapp’ application for iPad.
  • The field-based method for characterising PAWC has been tried and tested across Australia, but users need to be mindful of common pitfalls that can cause characterisation errors.
  • Knowledge of physical and chemical soil properties like texture or particle size distribution and (sub) soil constraints helps interpret the size and shape of the PAWC profiles of different soils. It can also assist in choosing a similar soil from the APSoil database.
  • Extrapolating from the point-based dataset to predict PAWC at other locations of interest is a challenge that needs further research. Preliminary analyses drawing on soil landscape mapping (NSW) and land resource area (LRA) mapping (Queensland) suggest that an understanding of position in the landscape and the story of its development may assist with extrapolation. This is because in many landscapes the soil properties determining PAWC are tightly linked to a soil's development and position in the landscape and these same aspects underpin soil and land resource surveys.
  • While the concept of using soil-landscape information to inform land management is not new (e.g. Queensland land management manuals draw on the same concept), the availability of these maps on-line makes them more accessible and assists with visualising a location’s position in the landscape. Combining these maps with the geo-referenced APSoil PAWC characterisations will increase the value that both resources can provide to farmers and advisors.
  • Uncertainty of PAWC estimates translates into uncertainty in PAW. The extent to which this affects potential decision making depends on the question asked, but also needs to be viewed in terms of the spatial variability in PAW and the accuracy of the method to convert this water into a yield forecast.

Plant available water and crop management decisions

A key determinant of potential yield in dryland agriculture is the amount of water available to the crop, either from rainfall or stored soil water. In the northern region the contribution of stored soil water to crop productivity for both winter and summer cropping has long been recognized. The amount of stored soil water influences decisions to crop or wait (for the next opportunity or long fallow), to sow earlier or later (and associated variety choice) and the input level of resources such as nitrogen fertiliser.

The amount of stored soil water available to a crop - Plant Available Water (PAW) – is affected by pre-season and in-season rainfall, infiltration, evaporation and transpiration. It also strongly depends on a soil’s Plant Available Water Capacity (PAWC), which is the total amount of water a soil can store and release to different crops. The PAWC, or ‘bucket size’, depends on the soil’s physical and chemical characteristics as well as the crop being grown.

Over the past 20 years, CSIRO in collaboration with state agencies, catchment management organisations, consultants and farmers has characterised more than 1000 sites around Australia for PAWC. The data are publicly available in the APSoil database, including via a Google Earth file and in the ‘SoilMapp’ application for iPad (see Resources section).

A number of farmers and advisers, especially in the southern Australia, are using the PAWC data in conjunction with Yield Prophet® to assist with crop management decisions. Yield Prophet® is a tool that interprets the predictions of the APSIM cropping systems model. It uses the information on PAWC along with information on pre-season soil moisture and mineral nitrogen, agronomic inputs and local climate data to forecast the possible yield outcomes at any time during the growing season. Yield Prophet® first simulates soil water and nitrogen dynamics as well as crop growth with the weather conditions experienced to date and then uses long term historical weather record to simulate what would have happened from this date onwards in each year of the climate record. The resulting range of expected yield outcomes can be compared with the expected outcomes of alternative varieties, time of sowing, topdressing, etc. to inform management decisions.

Others use the PAWC data more informally in conjunction with assessments of soil water (soil core, soil water monitoring device or depth of wet soil with a push probe) to estimate the amount of plant available water. Local rules of thumb are then used to inform the management decisions.

The APSoil database provides geo-referenced data (i.e. located on a map), but the PAWC characterisations are for points in the landscape. To use this information one needs to find a similar soil. This is not a straight forward process and subject of ongoing research, but a number of data and information sources are available that can assist. If suitable PAWC data are not found, local measurement of PAWC is required. This will often also provide a more accurate estimate although spatial variability may still be an issue.

This paper describes the measurement of PAWC, including practical tips and pitfalls, and outlines where to find existing information on PAWC. It discusses the principles behind extrapolation from known soil profiles and illustrates this with examples of PAWC data for local soils.

Plant Available Water Capacity (PAWC)

To characterise a soil’s PAWC, or ‘bucket size’, we need to determine (Figure 1a):

  • Drained upper limit (DUL) or field capacity – the amount of water a soil can hold against gravity
  • Crop lower limit (CLL) – the amount of water remaining after a particular crop has extracted all the water available to it from the soil
  • Bulk density (BD) – the density of the soil, which is required to convert measurements of gravimetric water content to volumetric water content.

In addition, soil chemical data are obtained to provide an indication whether subsoil constraints (e.g. salinity, sodicity, boron and aluminium) may affect a soil’s ability to store water, or the plant’s ability to extract water from the soil.

Figure 1. (a) The Plant Available Water Capacity (PAWC) is the total amount of water that each soil type can store and release to different crops and is defined by its Drained Upper Limit (DUL) and its crop specific crop lower limit (CLL); (b) Plant Available Water (PAW) represents the volume of water stored within the soil available to the plant at a point in time. It is defined by the difference between the current volumetric soil water content and the CLL.

Figure 1. (a) The Plant Available Water Capacity (PAWC) is the total amount of water that each soil type can store and release to different crops and is defined by its Drained Upper Limit (DUL) and its crop specific crop lower limit (CLL); (b) Plant Available Water (PAW) represents the volume of water stored within the soil available to the plant at a point in time. It is defined by the difference between the current volumetric soil water content and the CLL.

Plant Available Water (PAW)

Plant available water is the difference between the CLL and the volumetric soil water content (mm water / mm of soil) (Figure 1b). The latter can be assessed by soil coring (gravimetric moisture which is converted into a volumetric water content using the bulk density of the soil) or the use of soil water monitoring devices (requiring calibration in order to quantitatively report soil water content).

An approximate estimate of PAW can be obtained from knowledge of the PAWC (mm of available water/cm of soil depth down the profile) and the depth of wet soil (push probe or based on a feel of wet and dry limits using an uncalibrated soil water monitoring device).

Knowledge of PAW can inform management decisions and many in the northern region have, formally or informally, adopted this. Several papers at recent GRDC Updates have illustrated the impact of PAW at sowing on crop yield in the context of management decisions (see e.g. Routley 2010, Whish 2014, Dalgliesh 2014 and Fritsch and Wylie 2015).

Field Measurement of PAWC

Field measurement of DUL, CLL and BD are described in detail in the GRDC PAWC Booklet ‘Estimating plant available water capacity’ (see Resources section). Briefly, to determine the DUL an area of approximately 4 m x 4m is slowly wet up using drip tubing that has been laid out in spiral (see Figure 2). The area is covered with plastic to prevent evaporation and after the slow wetting up it is allowed to drain (see GRDC PAWC booklet for indicative rates of wetting up and drainage times). The soil is then sampled for soil moisture and bulk density.

The CLL is measured either opportunistically at the end of a very dry season or in an area protected by a rainout shelter between anthesis/flowering and time of sampling (Figure 2). This method assumes the crop will have explored all available soil water to the maximum extent and it accounts for any subsoil constraints that affect the plant’s ability to extract water from the soil.

Figure 2. Wetting up for DUL determination and rainout shelter used for CLL determination. Figure 2. Wetting up for DUL determination and rainout shelter used for CLL determination.

Figure 2. Wetting up for DUL determination and rainout shelter used for CLL determination.

Pitfalls and common mischaracterization issues

While the concept of PAWC is simple and the measurement methods for DUL and CLL were developed to be straightforward and not require any sophisticated equipment, it is important to keep an eye out for possible sampling errors. The list below summarises some of the key pitfalls and common mischaracterisation issues that we have come across in our collective experience of PAWC characterisations across Australia.

To allow interpretation and use of the data by others, PAWC characterisations should be accompanied by as much extra information as possible, including descriptions of the landscape position, surface condition (e.g. cracking, waterlogging), colour, texture (ideally with a full particle size analysis), Australian soil classification and any local classification soil name.

DUL

  • Weeds are often seen growing on the side of the plastic cover. It is important that these are strictly controlled throughout the wetting up process until sampling.
  • In sandy-textured soils the concentric rings of dripper line must be laid sufficiently close to each other to ensure consistent wetting across the whole area.
  • Allowing insufficient time for drainage may lead to overestimation of DUL, especially at depth. Heavier soils can take 1-2 months to drain.
  • Insufficient water application or application at too high a rate leads to underestimation of DUL at depth. This is particularly an issue with heavy clay soils, dispersive sodic soils and strong duplex (texture contrast) soils where water may move sideways. Both the GRDC PAWC booklet and the 'Soil Matters' book provide indicative rates and amounts for different soils. The wetting and drainage processes may be monitored (e.g. using NMM or a moisture probe), but this is not often done due to cost constraints (time, money).
  • Bulk density sampling, which is often done in conjunction with DUL sampling, requires a relatively high level of precision as any error in bulk density values will propagate when used to convert gravimetric water contents (including DUL, CLL and PAW) into mm of water. The procedure is described and illustrated in detail in the GRDC PAWC booklet.
  • Snakes like to hide under the plastic, so take care when wetting and sampling the plot.

CLL

  • The CLL method as described above relies on crop roots exploring the soil to the fullest extent. If the crop had insufficient moisture to establish its root system prior to anthesis, the CLL may not reflect maximum soil water extraction. Roots will not grow through a dry layer even if there is moisture underneath. It is, therefore, important to perform CLL measurement in paddocks with a well established and healthy crop. Wetting up of the CLL site prior to the growing season may help, but requires close attention to weeds and to supplying the right amount of nitrogen fertiliser.
  • In wetter climates and years with rainfall in the weeks just prior to the erection of rainout shelters at anthesis may refill the PAWC ‘bucket’. If the PAWC is large, this may prevent the crop from using all soil water and result in an overestimate of CLL (too wet). Ideally CLL is measured over multiple seasons, but this is rarely done in practice. Calibrated moisture probes can be an effective tool to assess a crop’s ability to extract moisture over a range of different seasons.
  • The CLL measured for one crop type may not apply to a different crop type, especially where growing season length or susceptibility to subsoil constraints differs. It is possible that long-season varieties may extract water from a greater depth than short season varieties because of more extensive root development, and hence result in different CLL.
  • If sampling is not deep enough to capture the full root zone, PAWC will be underestimated. In this case the CLL and DUL do not reach the same value at the bottom of the profile.
  • If there is insufficient wetting of the profile prior or during the growing season, the measured CLL may reflect the CLL of a previous crop. If the current crop has a shallower root system this could cause the PAWC to be overestimated. Wetting up of the CLL site prior to the season may help. Taking a soil core when the rainout shelter is installed and comparing values against those determined at the time of final sampling can assist with interpretation of the data.
  • Rainout shelters have blown loose or away on occasions, so it is important to secure the sides firmly into the soil.
  • For duplex soils located on hills slopes > 3-5% or soils at the break of slope, subsurface lateral flow can cause soil wetting despite the presence of a well constructed rain-out shelter. Keep an eye on late season rainfall and note any unusual wetness in samples collected.
  • Sampling after harvest when the soils are dry and hard, or have hard layers can be tricky. Digging a soil pit can be a better alternative than soil coring from the surface in these situations.

General

  • Soil variability may mean there is more than one PAWC profile within the paddock. Variability in depth of layers, e.g. texture contrast in duplex soils, can occur over small distances. This makes mixing replicates and selecting a “representative soil” difficult.
  • High soil variability can cause the DUL and CLL measurements to effectively be on different soils (even though they are usually only 2-3 m apart). It is essential to measure DUL and CLL on the same soil type. Yield or soil maps may assist in deciding where to sample.

Where to find existing information on PAWC

Characterisations of PAWC for more than 1000 soils across Australia have been collated in the APSoil database and are freely available to farmers, advisors and researchers. The database software and data can be downloaded from here. The characterisations can also be accessed via Google Earth (KML file from APSoil website) and in SoilMapp, an application for the iPad available from the App store. The yield forecasting tool Yield Prophet® also draws on this database.

In Google Earth the APSoil characterisation sites are marked by a shovel symbol (see Figure 3a), with information about the PAWC profile appearing in a pop-up box if one clicks on the site. The pop-up box also provides links to download the data in APSoil database or spreadsheet format.

In SoilMapp the APSoil sites are represented by green dots (see Figure 3b). Tapping on the map results in a pop-up that allows one to ‘discover’ nearby APSoil sites (tap green arrow) or other soil (survey) characterisations. The discovery screen then shows the PAWC characterisation as well as any other soil physical or chemical analysis data and available descriptive information.

Most of the PAWC data included in the APSoil database has been obtained through the field methodology outlined above, although for some soils estimates have been used for DUL or CLL. Some generic, estimated profiles are also available. While field measured profiles are mostly geo-referenced to the site of measurement (+/- accuracy of GPS unit), generic soils are identified with the nearest, or regional town.

The report 'PROFILE descriptions – District guidelines for managing soils in north-west NSW' by Daniells et al. (2002) provides PAWC characterisations for 17 soils in the region drawing on the same methodology. In addition, this report provides valuable soil descriptions for areas around Coonabarabran, Coonamble, Moree, Pilliga, and Walgett.

Figure 3. Access to geo-referenced soil PAWC characterisations of the APSoil database via (top) Google Earth and (bottom) SoilMapp (APSoil discovery screens as inserts).

Figure 3. Access to geo-referenced soil PAWC characterisations of the APSoil database via (top) Google Earth and (bottom) SoilMapp (APSoil discovery screens as inserts).

Figure 3. Access to geo-referenced soil PAWC characterisations of the APSoil database via (top) Google Earth and (bottom) SoilMapp (APSoil discovery screens as inserts).

Factors that influence PAWC

An important determinant of the PAWC is the soil’s texture. The particle size distribution of sand, silt and clay determines how much water and how tightly it is held. Clay particles are small (< 2 microns in size), but collectively have a larger surface area than sand particles occupying the same volume. This is important because water is held on the surface of soil particles which results in clay soils having the ability to hold more water than a sand. Because the spaces between the soil particles tend to be smaller in clays than in sands, plant roots have more difficulty accessing the space and the more tightly held water. This affects the amount of water a soil can hold against drainage (DUL) as well as how much of the water can be extracted by the crop (CLL).

The effect of texture on PAWC can be seen by comparing some of the APSoil characterisations from the northern region, as illustrated below (Figures 4-8). The soil’s structure and its chemistry and mineralogy affect PAWC as well. For example, subsoil sodicity may impede internal drainage and subsoil constraints such as salinity, sodicity, toxicity from aluminium or boron and extremely high density subsoil may limit root exploration, sometimes reducing the PAWC bucket significantly.

The CLL may differ for different crops due to differences in root density, root depth, crop demand and duration of crop growth. Some APSoil characterisations only determined the CLL for a single crop. The CLL for wheat, barley and oats are often considered the same and that of canola can be found to be similar as well, but care needs to be taken with such extrapolations as different tolerances for subsoil constraints can cause variation between crops.

A detailed explanation of the factors influencing PAWC is included in the ‘Soil Matters – Monitoring soil water and nutrients in dryland farming’ book, a pdf of which is available for free online (see Resources section).

North Star

The available APSoil PAWC characterisations in the region around North Star NSW fall roughly into two groups: lighter, texture contrast soils with smaller PAWC (Figure 4a) and vertosols characterised by larger PAWC, although dependent on texture (Figure 4c and d) and presence of subsoil constraints (Figure 4b, c and d). The PAWC differed depending on crop type due to differing responses to soil chemical conditions and growing season length (Figure 4b, c and d), although where differences are small they could be due to measurement error, including seasonal differences. Soil depth will impact on rooting depth and may also be an important factor determining the magnitude of PAWC, especially on the non-cracking clay soils.

The Profile Descriptions report provides descriptions and PAWC information for soils a bit further south in a region labelled in the report as ‘Moree East’ around Warialda. It describes the black vertosols that are non-sodic and non-saline as having PAWC up to 265 mm. The non-cracking soils (red and brown) in this area have PAWC between 100 and 215 mm, depending on texture and soil depth.

There are a number of APSoil characterisations to the east of this around the Newell Highway, which show varying PAWC. Factors like soil texture, depth of soil and subsoil constraints would play a role in determining the PAWC profiles, much like the examples shown in Figure 4.

Figure 4. Soils near North Star (see (a), (b), (c) and (d) below:

Figure 4. Soils near North Star (see (a), (b), (c) and (d) below:

Figure 4. Soils near North Star (see (a), (b), (c) and (d) below:

(a) Red chromosol (texture contrast soil) (APSoil 240) near North Star. The PAWC in the surface soil seems a bit high for a chromosol, which would normally show a shift and widening of the PAWC bucket. It could be that the texture change was not picked up if it was too shallow. Unfortunately particle size information was not available. Depth of texture change and subsoil chemistry will affect PAWC of the texture change soils.

(b) Black Vertosol (APSoil 237) near North Star with subsoil constraints limiting the PAWC (narrowing of the PAWC profile from 60 or 80 cm. Different crops respond differently to the constraints.

(c), (d) Clay content affects the PAWC of the vertosols as illustrated by (c) a Grey Vertosol - heavy Brigalow (Apsoil 101) near Tulloona) and (d) Grey Vertosol – Light Brigalow (APSoil 102) near Tulloona. These two soils were located within the same paddock and descriptions ‘heavy’ and ‘light’ may have had local relevance describing a texture difference, but particle size analysis is not available so this could not be confirmed.

Choosing an APSoil characterisation

As shown above, the soil PAWC can vary significantly. How do we choose the most appropriate APSoil characterisation, if we are not in the position to do a local field PAWC characterisation? This is still research in progress, but some guidance can already be provided.

  • The nearest APSoil may not be the most appropriate as its soil, parent material and landscape position could be quite different (cf. Figure 4)
  • Compare soil with descriptions of the APSoil sites (texture, colour, soil classification, chemical analysis). More recently collected APSoil characterisations include chemical analysis and particle size. As illustrated in Figure 4 both particle size and subsoil constraints strongly affect the PAWC.
  • Dig a hole (soil auger, soil core, backhoe trench, roadside bank or cutting); note surface features (cracking, hard setting), subsoil issues (salinity, sodicity, etc), rooting depth. This can assist with APSoil selection as well as adapting an APSoil profile to local conditions (e.g. if depth of texture change or rooting depth is different).
  • A measured sowing soil water profile (convert to volumetric) needs to ‘fit’ between CLL and DUL and can assist with APSoil selection (Figure 1b). If the measured (volumetric) water content profile is below CLL or above DUL then the texture of the soil does not match that of the chosen APSoil.
  • Opportunistic CLL (e.g. soil core following a dry finish; convert to volumetric) can be compared with CLL of APSoil characterisations.
  • Check for nearby soil survey characterisations (SoilMapp, Espade, Queensland Globe (see Resources section) and local soil reports) to help describe soils.
  • Draw on soil-landscape mapping (where available) to find APSoil sites in similar landscape positions (see below).
  • Native vegetation is often a useful indicator of soil type too and is indeed often included in information about soil-landscape, land resource area and land systems units.

Using soil –landscape information

In many landscapes the soil properties are tightly linked to a soil's development and position in the landscape and these same aspects underpin the many soil and land resource surveys that have been carried out over the years and that are increasingly becoming available on-line. Many of these present a mapping of so-called soil-landscape units that are based on a combination of geology, landscape features like slope and relief, vegetation and groups of soils. Effectively the distribution of soil types described by these maps and their mapping units descriptions are based on a landscape model or story. These descriptions, where available, can be used to interpret and potentially extrapolate APSoil characterisations.

In parts of NSW these soil-landscape units can be accessed through the ESpade tool (see Resources section), which delineates the units and provides a description and typical soil profiles for each unit (see Figure 5). In parts of Queensland, similar land resource area (LRA) mappings are used as part of land management manuals (see Figure 6). Where this information is available, it may be possible to use it to find an APSoil site in a similar landscape position as a first approximation of PAWC.

The concept of using soil-landscape information to classify and inform soil properties is not new. The Queensland land management manuals accompanying the LRA maps draw on the same concept as do the 'Glovebox Guide to Soil of the Macquarie-Bogan Flood Plain' by Hulme (2003) and several 'Soil Specific Management Guidelines for Sugarcane Production' in different sugarcane growing areas from northern NSW to northern Queensland (e.g. Wood et al 2003). The availability of these maps on-line makes them more accessible and assists with visualising a location’s position in the landscape. Combining these maps with the geo-referenced APSoil PAWC characterisations will increase the value that both resources can provide to farmers and advisors.

Using these resources to inform or even predict PAWC profiles is, however, still research in progress. In particular its predictive power and spatial accuracy still needs to be assessed as well as the required level of soil and landscape information. Not all areas within the northern region are covered by these soil-landscape maps and knowledge of (hydraulic properties of) soils within these areas varies too. Another resource that may proof useful in the future, but requires further testing for its use in predicting PAWC profiles, is the new Soil and Landscape Grid of Australia (see Resources section) which provides digital soil and landscape attribute predictions at a spatial resolution of 90 m x 90 m).

Figure 5. Example of soil-landscape mapping available for parts of NSW through ESpade. Mapping unit description is available through a pdf report and the landscape can be highlighted on the map.

Figure 5. Example of soil-landscape mapping available for parts of NSW through ESpade. Mapping unit description is available through a pdf report and the landscape can be highlighted on the map.

Figure 6. Section of Central Darling Downs with Land Resource Areas (LRA) delineated on Google Earth map with APSoil sites indicated. The accompanying description assisted in explaining the differences between some of the APSoil characterisations.

Figure 6. Section of Central Darling Downs with Land Resource Areas (LRA) delineated on Google Earth map with APSoil sites indicated. The accompanying description assisted in explaining the differences between some of the APSoil characterisations.

Local soil and landscape mapping information

North Star

ESpade does not provide on-line access to soil-landscape mapping in this area and neither is land systems mapping available online. SoilMapp provides a broad distinction of where the heavier vertosols and lighter texture change soils may be, but within these there will be local exceptions and variations.

The Profile Descriptions report provides descriptions and PAWC information for soils a bit further south in a region labelled in the report as ‘Moree East’.

Acknowledgements

The research undertaken as part of this project is made possible by the significant contributions of growers through both trial cooperation and the support of the GRDC, the authors would like to thank them for their continued support. We also gratefully acknowledge the contributions of CSIRO colleagues and many collaborators and farmers to the field PAWC characterisations. Their feedback also helped prepare the list of ‘tips and tricks’. The information on PAWC presented in this paper heavily draws on the work over many years by Neal Dalgliesh. Discussions with him and others, including with those involved with soil-landscape mapping in NSW (Neil McKenzie, Rob Banks, Brian Murphy and Neroli Brennan) were invaluable for the development of concepts and ideas presented in this paper. We thank Sean Murphy for access to the Profile Descriptions report for north-west NSW and its authors for the work involved with the PAWC characterisations and soil descriptions. Claire Yung provided assistance with the preparation of graphs.

Resources

APSoil, PAWC methodology and national information

APSoil database (includes link to Google Earth file)

SoilMapp (soil maps, soil characterisation, archive and APSoil sites): Apple IPad app available from App store; documentation

GRDC PAWC booklet

Soil Matters book

Soil and Landscape Grid of Australia

Yield Prophet®

NSW

ESpade (soil-landscape and land systems mapping and reports, reports on soil characterisation sites from various surveys)

Unpublished soil-landscape maps exist for: Nyngan, Walgett, Narromine, Narrabri, Gilgandra

Soil Profile Descriptions - District guidelines for managing soils in north-west NSW (Daniells et al. 2002)

Queensland

Land Management Manuals

Land Resource Area (LRA) maps: Google Earth files or via the Queensland Globe

References

Dalgliesh N (2014) Practical processes for better soil water management. GRDC Update paper.


Daniels I, Manning B, Pearce L (2002) Profile descriptions. District guidelines for managing soils in north-west NSW. NSW Agriculture.


Fritsch S, Wylie P (2015) Managing the yield gap to achieve your yield potential on the western Downs. GRDC Update paper.


Hulme P (2003) Glove-box guide to the soils of the Macquarie – Bogan floodplain, Sustainable Soils Management, Warren.


Routley R (2010) Water use efficiency – Optimizing farming systems performance and balancing fallow length and sowing 


Whish J (2014) Sorghum yield risk vs starting soil moisture. GRDC Update paper. 


Wood A, Schroeder B, Stewart B (2003) Soil Specific Management Guidelines for Sugarcane Production. CRC for Sustainable Sugar Production, Townsville.


Contact details

Kirsten Verburg
CSIRO Agriculture
Ph: (02) 6246 5954
Email: kirsten.verburg@csiro.au

Brett Cocks
CSIRO Agriculture
Ph: (07) 4688 1580
Email: brett.cocks@csiro.au

Tony Webster
CSIRO Agriculture
Ph: (07) 4059 5002
Email: tony.webster@csiro.au

GRDC Project Code: CSP00170,